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Endoplasmic Reticulum 3-ketoacyl-CoA thiolase, peroxisomal 3-hydroxy-3- methylglutaryl- coenzyme A reductase 1 Delta(14)- sterol reductase Hydroxymethylglutaryl- CoA synthase mevalonate kinase Phosphomevalonate kinase Diphosphomevalonate decarboxylase Isopentenyl- diphosphate Delta-isomerase Farnesyl pyrophosphate synthase Farnesyl pyrophosphate synthase Squalene synthase squalene monooxygenase Lanosterol synthase Lanosterol 14-alpha demethylase Delta(14)- sterol reductase Methylsterol monooxygenase Methylsterol monooxygenase Methylsterol monooxygenase C-3 sterol dehydrogenase 3-keto sterol reductase Methylsterol monooxygenase Methylsterol monooxygenase Methylsterol monooxygenase C-3 sterol dehydrogenase 3-keto sterol reductase Sterol 24-C- methyltransferase C-8 sterol isomerase C-5 sterol desaturase C-22 sterol desaturase C-8 sterol isomerase C-5 sterol desaturase Acetyl-CoA CoA Acetoacetyl-CoA 3-Hydroxy-3- methylglutaryl- CoA NADPH Mevalonic acid NADP CoA ergosta- 5,7,22,24(28)- tetraen-3-β- ol H + NADPH Ergosterol NADP 7- Dehydrodesmosterol Desmosterol Acetyl-CoA Acetoacetyl-CoA H 2 O CoA Mevalonic acid ATP Mevalonic acid-5P ADP ATP Mevalonic acid-5P ADP (S)-5- Diphosphomevalonic acid ATP ADP P i Isopentenyl pyrophosphate CO 2 Isopentenyl pyrophosphate Dimethylallylpyrophosphate Isopentenyl pyrophosphate Geranyl-PP PP i Geranyl-PP Isopentenyl pyrophosphate Farnesyl pyrophosphate PP i NADPH Squalene NADP PP i Squalene O 2 NADPH (S)-2,3- Epoxysqualene H 2 O NADP Lanosterin O 2 NADPH 4,4- Dimethylcholesta- 8,14,24- trienol Formic acid NADP H 2 O NADPH 4,4-Dimethyl- 5a-cholesta- 8,24-dien-3- b-ol NADP H + O 2 NADPH 4α- hydroxymethyl- 4β-methyl-5α- cholesta- 8,24-dien-3β- ol NADP H 2 O H + O 2 NADPH 4α-formyl-4β- methyl-5α- cholesta- 8,24-dien-3β- ol H 2 O NADP O 2 NADPH 4β-methyl-4α- carboxy- cholesta- 8,24-dien-3β- ol H 2 O NADP NADP 3-Keto-4- methylzymosterol CO 2 NADPH NADPH H + NADP 4a- Methylzymosterol H + O 2 NADPH H 2 O NADP 4α- hydroxymethyl- 5α-cholesta- 8,24-dien-3β- ol H + O 2 NADPH 4α-formyl-5α- cholesta- 8,24-dien-3β- ol H 2 O NADP O 2 NADPH 4α-carboxy- 5α-cholesta- 8,24-dien-3β- ol H 2 O NADP NADP 5α-cholesta- 8,24-dien-3- one CO 2 NADPH NADPH H + Zymosterol NADP S- Adenosylmethionine fecosterol S- Adenosylhomocysteine H + Episterol H + O 2 ferrocytochrome c 5,7,24(28)- Ergostatrienol H 2 O ferricytochrome c H + NADPH O 2 ergosta- 5,7,22,24(28)- tetraen-3-β- ol NADP H 2 O 5a-cholesta- 7,24-dien-3B- ol 7- Dehydrodesmosterol Cholesterol Stearidonoyl- CoA CoA CE(18:0)