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Unknown Unknown Phosphocarrier protein HPr Phosphocarrier protein HPr PTS system beta- glucoside- specific IIBCA components PTS system glucose- specific IIA component β-D-Glucose 6-phosphate α,α-Trehalose 6-phosphate H2O β-D- Glucopyranuronic acid ATP Glucose 6-phosphate ADP H+ α,α-trehalose Gram-Negative Cell Periplasm Outer Membrane Inner Membrane Cytoplasm Plasmid