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allantoate
transporter
(peroxisome)
allantoate
transporter
(ER)
Uricase
S-allantoin
synthase
S-allantoin
synthase
allantoinase
allantoate
amidohydrolase
ureidoglycine
aminohydrolase
ureidoglycolate
amidohydrolase
Uric acid
Allantoic acid
Allantoic acid
Allantoic acid
Allantoic acid
O
2
H
2
O
5-
Hydroxyisourate
H
2
O
2
H
2
O
5-hydroxy-2-
oxo-4-ureido-
2,5-dihydro-
1H-imidazole-
5-carboxylate
H
+
H
+
Allantoin
CO
2
H
2
O
H
+
H
+
H
2
O
S-ureidoglycine
Ammonium
CO
2
H
2
O
(S)-
Ureidoglycolic
acid
Ammonium
H
+
H
2
O
Glyoxylic acid
Ammonium
CO
2
Zinc (II) ion
Manganese
Manganese
purine
nucleotides
degradation
Peroxisome Membrane
Endoplasmic
ReticulumMembrane
Peroxisome
Endoplasmic Reticulum
Cytosol
Peroxisome
Endoplasmic Reticulum
?
?
At2g26230
TTL
TTL
ALN
AAH
UGLYAH
UAH
Uric acid
Allantoic acid
Allantoic acid
Allantoic acid
Allantoic acid
Oxygen
Water
5-
Hydroxyisourate
Hydrogen
peroxide
Water
5-hydroxy-2-
oxo-4-ureido-
2,5-dihydro-
1H-imidazole-
5-carboxylate
Hydrogen Ion
Hydrogen Ion
Allantoin
Carbon dioxide
Water
Hydrogen Ion
Hydrogen Ion
Water
S-ureidoglycine
Ammonium
Carbon dioxide
Water
(S)-
Ureidoglycolic
acid
Ammonium
Hydrogen Ion
Water
Glyoxylic acid
Ammonium
Carbon dioxide
purine
nucleotides
degradation
?
?
At2g26230
TTL
TTL
ALN
AAH
UGLYAH
UAH
Urate
Allanto
Allanto
Allanto
Allanto
O
2
H
2
O
5Hiso
H
2
O
2
H
2
O
5H2O4U2
H
+
H
+
Alantan
CO
2
H
2
O
H
+
H
+
H
2
O
Sug
Ammon
CO
2
H
2
O
Ureidog
Ammon
H
+
H
2
O
Glyoxal
Ammon
CO
2
Zinc
Mn2+
Mn2+
pu nu d
Peroxisome Membrane
Endoplasmic
ReticulumMembrane
Peroxisome
Endoplasmic Reticulum
Cytosol