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Carbamoyl-
phosphate
synthase
[ammonia],
mitochondrial
Carbamoyl-
phosphate
synthase
[ammonia],
mitochondrial
Carbamoyl-
phosphate
synthase
[ammonia],
mitochondrial
Dihydroorotate
dehydrogenase
(quinone),
mitochondrial
Unknown
Unknown
Soluble
calcium-
activated
nucleotidase 1
Cytidine/uridine
monophosphate
kinase 2
Nucleoside
diphosphate
kinase 6
Soluble
calcium-
activated
nucleotidase 1
GTP:AMP
phosphotransferase
AK3,
mitochondrial
Inosine
triphosphate
pyrophosphatase
CTP synthase 2
Soluble
calcium-
activated
nucleotidase 1
Nucleoside
diphosphate
kinase 6
Soluble
calcium-
activated
nucleotidase 1
Cytidine/uridine
monophosphate
kinase 2
5'-nucleotidase
domain-
containing
protein 2
Guanine
deaminase
5'-nucleotidase
domain-
containing
protein 2
Uridine
phosphorylase
Dihydropyrimidine
dehydrogenase
[NADP(+)]
Dihydropyrimidinase
Beta-
ureidopropionase
Ribonucleoside-
diphosphate
reductase large
subunit
Nucleoside
diphosphate
kinase 6
Cytidine/uridine
monophosphate
kinase 2
5'-nucleotidase
domain-
containing
protein 2
Uridine-
cytidine kinase
2
Cytidine
deaminase
Thymidine
phosphorylase
Deoxycytidylate
deaminase
Thymidine
kinase,
cytosolic
Thymidylate
synthase
Thymidylate
synthase
Soluble
calcium-
activated
nucleotidase 1
Nucleoside
diphosphate
kinase 6
Soluble
calcium-
activated
nucleotidase 1
5'-nucleotidase
domain-
containing
protein 2
Thymidine
kinase,
cytosolic
Thymidine
phosphorylase
Dihydropyrimidine
dehydrogenase
[NADP(+)]
Dihydropyrimidinase
Beta-
ureidopropionase
Deoxyuridine
5'-triphosphate
nucleotidohydrolase
Nucleoside
diphosphate
kinase 6
Thymidylate
synthase
Ribonucleoside-
diphosphate
reductase large
subunit
Uridine-
cytidine kinase
2
Beta-
ureidopropionase
Uridine triphosphate
Cytidine triphosphate
Uridine 5'-diphosphate
CDP
dCTP
Thymidine 5'-triphosphate
β-Alanine
L-Glutamine
Carbamoyl
phosphate
Phosphoribosyl pyrophosphate
3-
Aminoisobutanoic
acid
ATP
H
2
O
Hydrogen
carbonate
ADP
P
i
Ureidosuccinic
acid
L-Dihydroorotic
acid
Quinone
Accumulation
Orotic acid
Orotidylic acid
Uridine 5'-monophosphate
H
2
O
P
i
ATP
ADP
ADP
ATP
H
2
O
P
i
ADP
AMP
H
2
O
PP
i
ATP
ADP
P
i
dCTP
H
2
O
P
i
ADP
ATP
H
2
O
Cytidine monophosphate
P
i
ADP
ATP
H
2
O
Cytidine
P
i
H
2
O
Uridine
NH
3
H
2
O
P
i
P
i
Accumulation
Uracil
Ribose
1-phosphate
NADP
Dihydrouracil
NADPH
H
2
O
Ureidopropionic
acid
H
2
O
NH
3
CO
2
H
2
O
dCDP
ATP
ADP
dCMP
ATP
ADP
H
2
O
Deoxycytidine
P
i
ATP
ADP
H
2
O
Deoxyuridine
NH
3
P
i
Deoxyribose
1-phosphate
H
2
O
dUMP
NH
3
ATP
ADP
5,10-Methylene-THF
5-Thymidylic acid
Dihydrofolic acid
ATP
dTDP
ADP
H
2
O
P
i
ATP
ADP
H
2
O
P
i
H
2
O
Thymidine
P
i
ATP
ADP
P
i
Thymine