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Branched-chain
amino acid
transport
system 3
carrier protein
L-threonine
dehydratase
Probable
2-ketoarginine
decarboxylase
AruI
Unknown
Probable
2-ketoarginine
decarboxylase
AruI
Unknown
Probable
2-ketoarginine
decarboxylase
AruI
Acetolactate
synthase
isozyme 3 small
subunit
Ketol-acid
reductoisomerase
(NADP(+))
Dihydroxy-acid
dehydratase
Branched-
chain-amino-
acid
aminotransferase
High-affinity
branched-chain
amino acid
transport
ATP-binding
protein BraG
High-affinity
branched-chain
amino acid
transport
ATP-binding
protein BraF
High-affinity
branched-chain
amino acid
transport
system permease
protein BraD
High-affinity
branched-chain
amino acid
transport
system permease
protein BraE
Leucine-,
isoleucine-,
valine-,
threonine-, and
alanine-binding
protein
L-Threonine
L-Isoleucine
L-Isoleucine
Na
+
Na
+
H
+
H
2
O
(2Z)-2-
aminobut-2-
enoate
2-
iminobutanoate
H
2
O
H
+
Ammonium
2-Ketobutyric
acid
H
+
Pyruvic acid
CO
2
(S)-2-Aceto- 2-
hydroxybutanoic
acid
NADPH
H
+
NADP
(R) 2,3-
Dihydroxy-3-
methylvalerate
H
2
O
3-Methyl-2-
oxovaleric acid
L-Glutamic acid
Oxoglutaric
acid
2-Keto-3-
methyl-
valerate
L-Isoleucine
ATP
H
2
O
ADP
P
i
H
+
Threonine
Biosynthesis
braZ
ilvA
aruI
Unknown
aruI
Unknown
aruI
ilvH
ilvC
ilvD
ilvE
braG
braF
braD
braE
braC
L-Threonine
L-Isoleucine
L-Isoleucine
Sodium
Sodium
Hydrogen Ion
Water
(2Z)-2-
aminobut-2-
enoate
2-
iminobutanoate
Water
Hydrogen Ion
Ammonium
2-Ketobutyric
acid
Hydrogen Ion
Pyruvic acid
Carbon dioxide
(S)-2-Aceto- 2-
hydroxybutanoic
acid
NADPH
Hydrogen Ion
NADP
(R) 2,3-
Dihydroxy-3-
methylvalerate
Water
3-Methyl-2-
oxovaleric acid
L-Glutamic acid
Oxoglutaric
acid
2-Keto-3-
methyl-
valerate
L-Isoleucine
Adenosine
triphosphate
Water
Adenosine
diphosphate
Phosphate
Hydrogen Ion
Threonine
Biosynthesis